AFRC Veterinary Schools' Fellow, University of Cambridge: 1985-1989 MRC University Fellow, University of Oxford: 1989-1990 Veterinary Research Officer, AFRc/MRC Neuropathogenesis Unit, Edinburgh: 1990-1997 Senior Lecturer/Teacer, Departent of Veterinary Pathology, University of Glasgow: 1997-2003 Professor of Pathology & Infectious Diseases, Royal Veterinary College, London: 2003-2009 Professor of Veterinary Diagnostic Pathology, University of Cambridge: 2009-present